The Devonian Old Red Sandstone (ORS) magnafacies of southern Ireland is hosted in the Early Devonian Dingle Basin and the Late Devonian Munster Basin. Following the closure of the Iapetus Ocean during the Caledonian Orogeny, the Dingle Basin developed as a pull-apart structure before being deformed by Acadian tectonic activity. The Munster Basin developed as a half-graben structure in response to post-Acadian north–south extension in the region. Thus, the Irish ORS provides insights into the region’s tectonic history owing to its temporal and spatial proximity to the Caledonian (c. 475– 425 Ma), Acadian (c. 400–390 Ma) and Variscan orogenic events (c. 390–290 Ma). This study presents the first detrital zircon and apatite U–Pb geochronological data for the Upper ORS (UORS) in southern Ireland in addition to detrital white mica⁴⁰Ar/³⁹ Ar geochronological data to help unravel the depositional history of the Irish UORS and to assess the possible role of sedimentary recycling in Late Devonian basin development. Most UORS samples contain few late Neoproterozoic detrital zircon grains and are instead dominated by early Paleozoic and c. 1.1 Ga zircons. These populations represent recycling of northerly-derived Ordovician to Silurian strata of the Southern Uplands–Longford–Down terrane, which are of Laurentian affinity, and not recycling of Lower ORS (which contains a significant number of late Neoproterozoic detrital zircons) as previously thought. Similar detrital zircon dates have been observed in Givetian–Frasnian quartzites of the Pulo do Lobo Zone on the Iberian Peninsula, providing a possible Rheic Ocean link with the UORS.